Tanjiro joins the ranks of the Demon Slayers Corps to look for clues that will help return his half-demon sister Nezuko to human form. He boards the Infinity Train with Zenitsu and Inosuke and is joined by Rengoku, the Flame Pillar of the Demon Slayer Corps. As passengers on the speeding train start to disappear one by one without a trace, Rengoku senses that a demon is tormenting people. Tanjiro’s gang and Rengoku embark on a bloody struggle against the unfathomable, invincible demon to save the lives of the people on board.